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Chemolithotrophs
Oxidize INORGANIC compounds to obtain ENERGY
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Chemoorganotrophs
Oxidize ORGANIC compounds to obtain ENERGY
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Anaerobe
Uses CO2 as its carbon source
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Anaerobic chemolithotrophs
- Methanogens: OXIDIZE hydrogen gas to generate methane. Use CO2 as final e- acceptor
- Ie: Euryarchaeota
- Found in sewage, swamps, sensitive to oxygen
- Used in bioremediation
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Anaerobic chemoorganotrophs � anaerobic respiration
- Sulfur and sulfate reducing Bacteria: generate hydrogen SULFIDE, use Sulfate as final e- acceptor.
- Ie: Proteobacteria
- Gram NEG, Essential for sulfur cycle in ecosystem, responsible for rotten egg smell, found in organic matter
- Rxn ie: Organic comp + S (final e-acceptor) -> CO2 + H2S (hydrogen sulfide)
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Anaerobic chemoorganotrophs � fermentative
- ALL GRAM +
- Clostridium: endospore forming obligate anaerobes
- Lactic Acid Bacteria � Streptococcus: Lactic acid is major end product of Fermentation, energy only comes from Substrate level phosphorylation. Most can grow in aerobic environments
- Ie for both above: FirmiCUTES
- Propionibacterium: Propionic acid is major end product of fermentation. Swiss cheese
- Ie: Actinobacteria
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Earliest organism of Oxygenic phototrophs
Cyanobacteria: act to harvest sunlight to produce organic compounds thru conversion of CO2
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Anoxygenic phototrophs
OXIDIZE hydrogen sulfide or ORGANIC molecules when making NADPH
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Oxygenic phototrophs
Photosynthetic bacteria that use WATER as source of electrons. OXIDATION of H20 liberates Oxygen
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Anoxygenic phototrophs
- ALL Gram NEGATIVE
- Purple Sulfur bacteria (ie Chromatium): Use SULFUR compounds (prefer hydrogen sulfide) as a source of electrons when making reducing power. Ie- Proteobacteria
- Purple Non-sulfur Bacteria (ie Rhodobacter): Use ORGANIC compounds as source of Electrons for reducing power. Found in soil, diverse metabolism. Ie- Proteobacteria
- Green Sulfur bacteria (ie: Chlorobium): Found in habitats similar to purple sulfur bac. ALL are strict ANAEROBES. Ie- Chlorobi
- Chloroflexus: Characterized by filamentous growth. Ie- Chloroflexi
- Others: Heliobacterium- ie-firmicutes
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Oxygenic phototrophs
- Nitrogen-fixing cyanobacteria: incorporate Nitrogen gas & CO2 into organic material to produce ammonia. Helps control CO2 atmosphere levels.
- Occurs in HETEROCYST (protects breakdown of nitrogenase from oxygen)
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Aerobic chemolithotrophs
- Obtain energy by OXIDIZING reduced inorganic chemicals.
- These use oxygen as the final e- acceptor
- Include sulfur-oxidizing bacteria, nitrifiers and hydrogen-oxidizing bacteria
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Aerobic chemoorganotrophs
- OXIDIZE organic compounds to obtain energy
- Use oxygen as final e- acceptor.
- Can be classified as: Obligate aerobes or facultative anaerobes
- Ie: Organic compound + O2 -> CO2 + H20
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Aerobic chemolithotrophs
- ALL Gram NEGATIVE
- All seen with Proteobacteria (exceptHYDROGEN oxidizing bacteria, ie is Aquafacae)
- Sulfur-Oxidizing bacteria (ie- Beggiatoa): OXIDIZE reduced sulfur components, sulfuric acid is produced. Ie- S + O2 -> H2SO4 + H20. Live in sulfur springs, cause bulking in sewage treatment.
- Unicellular Sulfur Oxidizers (Thiobacillus): same as above, but some produce enough acid to lower pH to 1.0.
- Nitrifiers (ie- Nitrococcus): OXIDIZE inorganic nitrogen (such as ammoniam(NH4) or Nitrite(NO2)) to obtain ENERGY. Ie- NH4 + O2 -> NO2 + H20 + H
- These deplete O2 in waters that are contaminated with ammonia waste
- Hydrogen Oxidizing Bacteria (ie- Aquifex): Obligate Chemolithotrophs, found in HOT springs. Ie- H2 + O2 -> H20
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Aerobic chemoorganotrophs (OBLIGATE Aerobes)
- OBLIGATE Aerobes (ONLY Aerobic respiration)
- Micrococcus: Gram + in soil. Ie- Actinobacteria
- Mycobacterium (TB): Gram +, lives on DEAD or decaying matter. Ie- Actinobacteria
- Pseudomonas: Gram Neg, motile and pigmented, opportunistic. Ie- Proteobacteria
- Thermus: produces Taq. Ie- Deinococcus-Thermus
- Deinococcus: used to cleanup Radioactive contamination Ie- Deinococcus-Thermus
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Aerobic chemoorganotrophs (Facultative Anaerobes)
- Corynebacterium: Gram + rods, inhabits soil, H20. Ie- Actinobacteria
- Enterobacteriaceae (ie- E. coli): Gram neg rods, live in intestinal tract. Ie- Proteobacteria
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Example of a medically important terrestrial thriving chemoorganotroph
Argobacterium: causes plant tumors, used by scientists to genetically modify plants to have higher crop yields. ie- (phylum is...Proteobacteria)
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Example of a medically important aquatic thriving chemoorganotroph
Sheathed bacteria (Sphaerotilus): forms chains of cells enclosed within a protective sheath. ie- proteobacteria
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List the kinds of bacteria that Form endospores
Clostridium (anaerobic chemoorganotroph): is a GRAM + rod
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List and describe the kinds of prokaryotic microorganisms that live in an aquatic environment.
- Sulfur Oxidizing bacteria, Nitrifiers & hydrogen oxidizing bacteria (aerobic chemolithotroph)
- Purple Sulfur and Non-sulfur bacteria (anoxygenic phototroph)
- Methanogens (anaerobic chemolithotrophs)
- Corynebacterium (aerobic chemoorganotrophs)
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List and describe the kinds of prokaryotic microorganisms that use animals as their habitat.
- Methanogens: found in the digestive tract of mammal (anaerobic chemolithotrophs)
- Enterobacteriaceae (ie- E. coli): Gram neg rods, live in intestinal tract
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List and describe the kinds of prokaryotic microorganisms that live under extreme conditions.
Hydrogen-oxidizing bacteria (aerobic chemolithotrophs) can survive and thrive at 95 Celsius.
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