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Hemichordates
- means "partial chord" (acorn worm and pterobranchs)
- are worm like marine animals
- range from 2 cm to 2 m
- filter feeders
- sedentary
- live under stones, in burrow, or in tubes if in shallow water
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why are hemi chordates like chordates
more like chordates as they have ciliated pharengeal slits, an open cirulatory system, and a dorsal hollow nerve tube in the collar region
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Chordates
- name chordata comes from: the noto chord which is a rod like semi rigid organ enclosed in a sheath
- they posess: a dorsal hollow nerve cord extending the length of their body, pharyngeal slits, an endostyle (ciliated groove in the pharynx) and a mucular post anal tail.
- 3 sub phyla: urochordata, cephalochordata, craniata
- latter includes infra phyla hyperotreti and vertebrata
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Subphylum Urochodata (Tunicates)
- 3000 spp
- sessile
- aquatic
- only larval form has all chordate characteristics including notochord
- body wall made of connective tissue called a tunic
- have a highly developed muscular pharyngeal basket used in filter feeding
- larvae are beleived to to be the prototype of ancestral chordate
- chordates evolved from this via paedomorphisis (sexual maturation of the larval form)
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Subphylum Cephalochordata (head chord)
have well developed notochord through out their life, gill bars, segmented muscles, fish shapedbody but no vertebrae, brain or anterior sensory organs
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Lanclets- Amphioxus
- small (5-8cm)
- latterally flattened
- transluscent
- live in sandy bottoms of temperate costal waters
- extract food from sea water by filter feeding while burrowed in the sand with only their head exposed
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Infraphylum hyperotreti
- includes many extinct fish like animals as well as the hag fishes
- have a fully developed notochord and a cartelagenous skull but no vertebrae or jaws
- used to be group with lamp reys
- extant member hag fishes (class myxini) are fish like marine animals that burro in sand and mud
- they eat the inside of dead or dying fish by entering through an organisms mouth for defence they secret copious amounts of slims through special slime glands
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Inphraphylum Vertebrata
- like other chordates as they have gill bars and segmented muscles
- have modifications allowing for increase size speed and mobility
- defining feature is there segmented vertebral column that houses the dorsal nerve chord
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Lampreys
- jawless fish like animals like hag fish but unlike hag fish they have simple vertebrae
- larvae- mall bottom dwelling filterfeeders
- adults parasites on other fish
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Anatomy of lampreys
- 1horny teeth ----7intestine -----13respiratory tube
- 2buccal funnel --8liver----------14 velum
- 3mouth----------9gonad--------15olfactory sac
- 4tongue---------10notochord---16brain
- 5nostril----------11gill filamen---17 spinal chord
- 6pineal eye------12heart
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Integument
- lamprey skin is mooth and pliable, coverd with mucous
- epidermis is a living tissue of stacked layers of epidermal cells with unicellular glands (not normally kerotinized)
- dermis is in regular layers of collagenous connective tissue with pigment cells
- hypodermis includes adipose tissue
Question 1: observe the slide of the lamprey on the side bench and identify these features( epidermal cells, collageneous tissue, adipose tissue)
Answer 1: no scale on a lamprey
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Support and locomotion (the endo skeleton)
- secreted and maintained by living tissue
- expand as we grow
- superior to the exoskeleton for those over 0.5 grams
- early on skeletal system made of cartelage
- chondrichthyes (sharks and rays)skeleton is always cartilage. this is a derived feature as ancestors didnt have it
- functions: skeltetal support during development, cushioning between joints, absorption and storage of mechanical energy during locomotion.
- consists primarily of chondrocytes within a matrix of collogen
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types of bone
- endochondral bone: develops within the cartilage from osteocytes that have inveded the cartilage (most os the skeleton)
- dermal bone: developes beneath the skin. first type of bone, seen first in ostracoderms (extinct jawless fish that carried bony armour of dermal plates
- face and cranium made of dermal bone
Question 2: how do cartilage and bone differ at the microscopic level
- Answer 2:
- cartilage: chondrocyte cell in a mixture of collagen
- bone: bone matrix deposited in concentric layers around osteonic canals
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Muscles
- composed of muscle fibres comprised of thousands of myofibrils
- striated muscle: (voluntary and multi nucleate) used for gross movements such as locomotion. can also maintain circulation and regulate body temperature
- Smooth muscle: (involuntary and uninucleate) occurs in the skin and the lining of the gut, blood vessels, and reproductive tracts
- Cardiac muscle: occurs in the heart and requires no innervation are branched
Question 3: how do the types of muscle differ in their microscopic structure?
- skeletal: long cilindrycal shape
- smooth: spindle shaped
- cardiac: striated and branched
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Food aquisition and digestion
- the major function of the mouth is to aquire food
- also used for taste, digestion, defence, and aggression
- the mouth and the pharynx function in vocalization
Question 4: does the lampreyhave all teh major components of a vertebrate digestive system (teeth, tongue, salivary glands, stomach, liver, and intestines)? does this have andy functional significance (think in terms of lampreys diet-blood)?
- they have a protrusable toung to suck the blood out of their prey
- have many rows of teeth to grasp the sides of fish
- have 1 pair of salivar glands which have anti coagulates to keep sucking the blood of prey
- short esophagus because they suck blood
- contain a stomach (ancestral vertebrate structure). stomach allows for storage of large feedings of larger animals or large blood feedings
- do not have a gizzards as they dont grind up food
- have intestines to absorb blood
- dont have a ceacum
- dont have a liver or galbladder because the blood is already filtered
- have a pancrease to secrete hormones
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Gas Exchange and water balance
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Circulation
- circulatory system supplies nutrients and gases to body as well as removes wastes and excess heat (specialization necessary for those larger than a few millimeters in size)
- also helps in communication for cells (important for growth differentiation, and reproduction)
- blood is made up of: plasma, erythrocytes, and leukocytes
- Question 5: in what ways do the blood cells vary between the groups?
- Answer 5: they vary in size and nucleation (RBCs lack a nucleus)
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Excretion and osmoregulation
- The Kindey: balances salts and water (osmoregulation)
- process of digestionand respiration result in the accumulation of wasts which must be removed as they are lethal. removal of these wastes involves respiration (CO2) and excretion
- most vert tissue is 1% salt content compared to that of sea water
- hag fish evolution was likely entirely marine
- basic units of the kidney: the renal corpuscle, glomerulus, renal tube
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Pronephros
- most primitive kidney
- functional only in embryosof fishes and amphibians and in adult hagfishes (who are iso osmoticand secrete ammonia)
- contains the fewest nephrons
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Mesonephros
- functional kidney during embryonic developement in amniotes (reptiles birds and mammals)
- has more nephrons than the pronephros
- in adult fishes and amphibians kidney has mesonephric and meta nephric portions. this is called opsithonephric kidney
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Metanephros
- functional kidney of adult reptiles birds and mammals
- helps with water consevation
- largest number of nephrons and is the most effiecient
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Excrition
- ammonia: highly solluble, occurs in aquatic animals
- urea: less soluble, requires some energy
- uric acid: very low solubility, but requires even more energy (used by those in very dry areas)
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Sensory system
- percieves the external environment and transmit the intormation to th eorganism
- they participate in the regulation of the internal environment
- the nervouse system serves as a storage site for information
- nervous system is based on the elaboration of a dorsal nerve cord
- nerves are aggregations of nerve axons
- they seperate into dorsal (sensory) and ventral (motor) roots or horns
- consists of CNS, Peripheral nerves (spinal and cranial), and autonomic (sympathetic and parasympathetic)
- vertebrate brains is tripartite (3 lobes). 2 lobes divide to eventually give 5 lobes
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Neurons
- Sensory Neurons: carry info from a receptor to the CNS
- Motor Neurons: carry info from CNS to an effector
- Inter Neurons: co-ordinates sensory and motor neurons
- Consist of: nerve body: dendrites, axon
- Question 6: where are the cranial and spinal nerves located?
- cranial nerves are in the head
- spinal nerves are in the spinal chord
- Question 7: where do the nerves of the sympathetic system generally originate?
- at the thoracic and lumbar regions of the spinal chord
- Question 8: where do the nerves of the parasympathetic system generally originate?
- the brain and sacral region of the spinal chord
- Question 9: what are the primary functions of the 5 lobes of the brain?
- telencephalon: cerebrum division for specialization, thalamus sensory relay center, hypothalamus, hormones, and homeostasis
- diencephalon
- mesencephalon optic lobe optic reflex center, cerebellum- motor activity, pons- nerve tracks, breathing
- metencephalon medulla oblongata- reflex centers
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Sensory Structures
- 4 types
- Chemoreceptors: smell and taste
- Mechanoreceptors: touch hearing and pressure
- Photoreceptors: light
- Thermoreceptors: temperature
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Photoreception:
- Question 10:what are the functions of the lens, iris, pupil, and retina?
- lense- focuses incoming light onto the retina
- iris- alters the light intensity by controlling the size of the pupil
- pupil- the whole where the light enters
- retina- light sensitive membrans processes images
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Mechanoreception:
- hair cells convey information pertaing to posture accelleration, and sound
- statocysts of inner ear detect orientation and movement (semicircular canals and otolith organs (utriculus, and sacculus)
- similar mechanism lets fish detect movement in the water
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