-
Parsimony
(selection of trees)
The tree that is the simplest explanation of relationships is the tree adopted.
-
Maximum Likelihood
(selection of trees)
Molecules
-
Monopyletic
A group that includes the ancestrial species and ALL of its descendents
-
Paraphyletic
A group that consists of an ancestrial species, and some, but not all of the descendents
-
Polyphyletic
A grouping of several species that lack a common ancester.
-
Early Earth
4.6 billion years ago
Molten Planet
-
First Rocks
3.8 billion years ago
- 1. Abiotic synthesis of organic molecules
- 2. Synthesis of polymers
- 3. Protobionts
- 4. First Genetic Material
-
Abiotic (non-living) Synthesis of Organic Molecules = Abiogenesis
-1920's Oparin-Haldane Hypothesis
-1953 Stanley Miller & Harold Urey
-
Oparin-Haldane Hypothesis
- Reducing atmosphere + lightening/UV light =
- organic molecules
-
Stanley Miller & Harold Urey
- -created early earth's atmosphere (CH4 NH4 H2O H2S)
- -when they added lightening--->produced small organic molecules
- -this hypothesis was controversial since some support the idea that early earth was not very reducing & made up mainly of CO2 & N2 gas
-
Synthesis of Polymers
-replication by dripping solutions of amino acids on hot sand, clay or rock
-
Protobionts
Abiotic groups of molecules surrounded by a membrane
- ex: Liposomes = lipids + H2O
- -"replicate"
-
First Genetic Material
-Probably RNA, not DNA
-
Fossil Record - Provides the history of life
1. Relative Dating
2. Absolute Dating
-
Relative Dating
Certain rock layers above/below others w/ certain fossils give us a sequence of events
-
Absolute Dating
Determination of how old something is in calendar years
- -based on decay of radioactive isotopes
- -each radioactive isotope has half-life
ex: C-14 half life: 5,730 years
-
First Fossils
- Stromatolites = Bacterial Mats 3.5 bya
- -cyanobacteria
-
2.7-2.2 Billion years Ago
- O2 levels begin to creep up
- -banded-iron formations
-
2.1 Billion years ago
- First Eukaryotes
- -endosymbiont origin of mitochondria & plastids
- -mitochondria = aerobic heterotrophic prokaryote
- -plastid = photosynthetic prokaryote
- -serial endosymbiosis
-
Serial Endosymbiosis
- 1. Mitochondria endosymbiosis likely occured first
- -Found in all plants and animals
- 2. Plastids - only found in plants
- -Likely occured after mitochondrial endosymbiotic
- event
-
Evidence for Endosymbiosis
1. Mitochondria & plastids have their own DNA
- 2. Membranes of both are similar to those seen in
- prokaryotes
-
1.5 Billion years ago
- First Multicellularity
- -small in size, limited in distribution
- -likely evolved independently several times
- -not much advancement after this point for a long time
-
570-550 Million years ago
Larger, soft bodied organisms
-
Cambrian Explosion
- ~542 million years ago
- -first hard-shelled organisms
- -in first 20 million years most of the major animal phyla we know of originated (exceptions: Porifera-sponges & Chidaria-jellyfish)
- Reasons for this:
- 1. O2 levels
- 2. Predator-prey relationships
-
Archaean (Eon)
4.6 bya - 2.5 bya
-
Proterozoic (Eons)
2.5 bya - 542 mya
-
Phanerozoic (Eons)
542 mya - Present
-
Paleozoic Era
542 mya - 251 mya (age of invertebrates)
-
Mesozoic Era
251 mya - 65 mya (age of reptiles)
-
Cenozoic Era
65 mya - Present (age of mamals)
-
Major Events that Shaped Diversity During Phanerozoic
- 1. Plate techtonics
- 2. Mass Extinctions & Explosive Diversifications
-
Plate Techtonics
- -continents moving around via plate techtonics
- -come together, break apart, come together, etc.
- -create mountains, oceans, volcanoes, earthquakes
ex: Pangea
-
Pangea (250 mya)
- -a single, large super continent
- -reduced coastline
- -higher gene flow possible
- -more of the continental land mass is drier (more inland area)
- -ocean currents change
-
Mass Extinctions & Explosive Diversifications
End Permian Extinction
- -(251 mya)
- *largest extinction in Earths history
- *90% of all species go extinct
-
Mass Extinctions & Explosive Diversifications
End Cretaceous K-T Extinction
- -65 mya
- *asteroid impact in the Yucatan Peninsula (Alvarez,
- 1980)
- *wiping out dinosaurs (except birds)
-
Prokaryotes
2 Domains: Bacteria & Archaea
Features:
- 1. Smaller than Eukaryotes (1-5um)
- 2. Generally single celled
- 3. 3 common shapes (cocci, bacillus, spiral)
- 4. Cell wall present
- a) Bacteria-primarily PG
- b) Archaea-mainly polysaccharides & protein, no
- PG
- 5. Many cell walls covered by a capsule (sticky outer layer that helps organism adhere to substrates
- 6. About half can move in a direction (flagella)
- 7. Generally lack internal membrane bound structures
- 8. No nucleus
- 9. Reproduce via binary fission
- 10. Many have plasmids (smaller rings of DNA)
-
Autotrophs
Obtain food w/o eating other organisms
- A) Photoautotrophs-light, CO2
- B) Chemoautotrophs-inorganic chemicals, CO2
-
Heterotrophs
Obtain food by eating other organisms
- A) Photoheterotrophs-light, organic compounds
- B) Chemoautotrophs-organic compounds
-
Oxygen Metabolism
Obligate Aerobes
Use and need O2 for cellular respiration
-
Oxygen Metabolism
Facultative Aerobes
Use O2 if present, but can also use fermentation
-
Oxygen Metabolism
Obligate Anaerobes
Poisoned by O2, only fermentation or other chemical energy used
-
Nitrogen Metabolism
Various types of Nitrogen used
-
Bacteria-Proteobacteria
- Gram Neg.
- ex: E. Coli, Salmonella
-
Bacteria-Chlamydias
Gram neg., No PG
-
Bacteria-Spirochetes
- Helical
- ex: syphilis, lime disease
-
Bacteria-Gram +
Ex: leprasy, lime disease
-
Bacteria-Cyanobacteria
Photoautotrophs
-
Archaea
Live in extreme environments
- Thermophiles-hot environments
- Halophiles-Salty environments
- Methanogens-use CO2 and generate methane
-
Archaea-Synampomorphies shared w/ Eukarya
- 1. Lack PG in cell walls
- 2. Several kinds of RNA Polymerase
- 3. Introns present in DNA
- 4. Histones present (proteins associated w/ chromatin)
- 5. Use Methionine as initiator amino acid for protein synthesis
- 6. Growth not inhibited by antibiotics
|
|