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speciation
the process by which one species splits into two or more species; responsible for diversity of life
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microevolution
changes over time in allele frequencies in a population
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macroevolution
the broad pattern of evolution over long time spans
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biological species concept
defines a species as members of populations that actually or potentially interbreed in nature, not according to similarity of appearance; appearance is helpful in identifying species, but it doesn't define species; species are identified by the ABSENCE of gene flow
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reproductive isolation
prevents the formation of new species because of the existence of biological factors (barriers) that impeded members of two species from producing viable, fertile offspring; block gene flow between species and the formation of hybrids
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prezygotic barriers
block fertilization from occurring; (1) impede members of different species from attempting to mate (2) preventing an attempted mating from being completed successfully (3) hindering fertilization if mating is completed successfully
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postzygotic barriers
contribute to reproductive isolation once the zygote is formed; ex: developmental errors may reduce survival/problems after birth leave offspring infertile or unable to survive long enough to reproduce
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habitat isolation
2 species that occupy different habitats wihtin the same area may see each other rarely even if they aren't separated by geographic barriers; pre
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temporal isolation
species that breed during different times of day, seasons, or years can't mix gametes; pre
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behavioral isolation
courtship rituals that attract mates and other behaviors unique to a species are reproductive barriers; such rituals allow for mate recognition (a way to identify potential mates of the same species); pre
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mechanical isolation
mating is attempted, but morphological differences prevent its successful completion; pre
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gametic isolation
sperm of one species may not be able to fertilize the eggs of another species (ex: it cannot survive in reproductive tract of female); pre
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reduced hybrid viability
the genes of different parent species may interact in ways that impair the hybrid's development or survival in its environment; post
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reduced hybrid fertility
when hybrids are sterile; meiosis can fail to produce normal gametes if the two parent species differ in chromosome number or structure; post
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hybrid breakdown
when 1st generation hybrids are fertile and stable, but their offspring with either one another or a parent are feeble or sterile (steeble); post
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morphological species concept
characterizes a species by body shape and other structural features; can be applied to both asexual and sexual organisms
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ecological species concept
views a species in terms of its ecological niche
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phylogenetic species concept
defines a species as the smallest group of individuals that share a common ancestor, forming one branch on the tree of life
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allopatric speciation
gene flow is interrupted when a population is divided into geographically isolated subpopulations
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sympatric speciation
speciation occurs in populations that live in the same geographic area; less common than allopatric; can occur if gene flow is reduced by factors such as polyploidy, habitat differentiation, and sexual selection
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autopolyploid
an individual that has more than two chromosome sets that are all derived from a single species
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allopolyploid
an organism that has two sets of chromosomes, each from one parent; are usually only fertile when reproduction is asexual
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hybrid zone
a region in which members of different species meet and mate, producing at least some offspring of mixed ancestry
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reinforcement
when natural selection strengthens prezygotic barriers because hybrids are less fit than members of their parent species; formation of hybrids is reduced; because of this, barriers of reproduction between species should be stronger for sympatric species than for allopatric species
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punctuated equilibria
periods of apparent stasis punctuated by sudden change
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phylogony
the evolutionary history of a species or group of species
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systematics
a discipline focused on classifying organisms and determining their evolutionary relationships; used to construct phylogenies
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binomial
the first part of the binomial naming system is the genus and is where the species belongs; the second part is called the specific epithet and is unique for each species within the genus
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taxon
the named taxonomic unit at any level of the hierarchy
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phylogenetic tree
the evolutionary history of a group of organisms can be represented in a branching diagram
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PhyloCode
a classification based entirely on evolutionary relationships; it only names groups that include a common ancestor and all of its descendants
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sister taxa
groups of organisms that share an immediate common ancestor and are therefore each other's closest relatives
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a tree that is rooted...
means that a branch point within the tree represents the last common ancestor of all taxa in the tree
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polytomy
a branch point from which more than two decendant groups emerge; indicates that evolutionary relationships among the descendant taxa are not yet clear
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analogy
a similarity due to convergent evolution; a bat wing is ANALOGOUS to a bird's wing
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homoplasies
analogous structures that arose independently (from the Greek "to mold in the same way")
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molecular homoplasies
shared nucleotide bases in organisms that are otherwise very different; coincidental matches
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molecular systematics
uses DNA and other molecular data to determine evolutionary relationships
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cladistics
biologists place species into groups called clades, each of which includes an ancestral species and all of its decendents
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monophyletic
a taxon (group of organisms) which forms a clade, meaning that it contains all the descendants of the possibly hypothetical closest common ancestor of the members of the group
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paraphyletic
a group which consists of an ancestral species and some, but not all, of its decendents
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polyphyletic
includes taxa with different ancestors
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shared ancestral character
a character that originated in an ancestor of the taxon; (ex: in mammals, the backbone)
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shared derived character
an evolutionary novelty unique to a particular clade (ex: in mammals, hair)
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outgroup
a species or group of species from an evolutionary lineage that is known to have diverged before the lineage that includes the species we are studying (ingroup)
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maximum parsimony (Occam's Razor)
it is important to first investigate the simplest explanation that is consistent with the facts
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maximum likelihood
given certain rules about how DNA changes over time, a tree can be found that reflects the most likely sequence of evolutionary events
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phylogenetic bracketing
we can predict (using parsimony) that features shared by two groups of closely related organisms are present in their common ancestor and all of its descendants
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orthologous genes
homologous genes that are found in different species because of speciation; widespread and can extend over huge evolutionary distances
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paralogous genes
result from gene duplication, so they are found in more than one copy in the same genome
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molecular clock
a yardstick for measuring the absolute time of evolutionary change baseed on the observation that some genes and other regions of genomes appear to evolve at constant rates
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neutral theory
much of evolutionary change in genes and proteins ahs no effect on fitness and therefore is not influenced by Darwinian selection
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horizontal gene transfer
the movement of genes between organisms in different domains; the process in which genes are transfered form one genome to another through mechanisms such as exchange of transposable elements and plasmids, viral infection, and PERHAPS fusion of organisms
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