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evolution
descent with modification; change in the genetic composition of a population from generation to generation
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Aristotle and Evolution
viewed species as unchanging; thought that life-forms could be arranged on a ladder of increasing complexity = SCALA NATURAE
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Linnaeus
developed the binomial (two-part) system of naming species; was a nested system of classification grouping similar species into increasingly general categories
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fossils
remains/traces of organisms form the past; most are found in sedimentary rocks formed from sand and mud that settled to the bottom of bodies of water
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catastrophism
the idea that past events occurred suddenly and were caused by mechanisms DIFFERENT from those operating in the present; advocated by Cuvier, who speculated that each boundary between strata indicated the occurence of a catastrophe
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uniformitarianism
idea that mechanisms of change are constant over time; Lyell (using Hutton) speculated that the same geological processes are operating today as in the past AND at the same rate
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Lamarck
found lines of decent and explained them using use and disuse (parts of the body that are used become larger/stronger while those that go unused deteriorate) AND inheritence of acquired characteristcs (an organism could pass these modifications to its offspring)
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adaptations
chaaceristics of organisms that enhance their survival and reproduction in specific environments
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natural selection
individuals with certain inherited traits leave more offspring that individuals with other traits; a process of editing rather than a creative mechanism; doesn't create but SELECTS for individuals already present in a population
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Darwin developed two ideas in his book:
- 1) descent with modification explains life's unity & diversity
- 2) natural selection brings about the match between organisms and their environment
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artificial selection
when humans modify other species over many generations by selecting and breeding individuals that possess desired traits; such organisms often bear little resemblance to their wild ancestors
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cetaceans
mammalian order that includes whales, dolphins and porpoises
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homology
similarities between species with characteristics that have underlying similarities even thought they might have very different functions; a result of common ancestry
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homologous structures
variations on a structural theme that was present in their common ancestor; ex: anatomical resemblances in the skeletons of arms, forelegs and flippers of mammals
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vestigal structures
remnants of features that served important functions in the organism's ancestors
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evolutionary tree
a diagram that reflects evolutionary relationships among groups of organisms
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convergent evolution
the independent evolution of similar features in different lineages; responsible for analagies
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analogous
features in different species' that have adapted to similar environments in similar ways; resemblances due to species sharing features because of convergent evolution
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biogeography
the geographic distribution of species; influenced - among other things - by continental drift
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continental drift
the slow movement of Earth's continents over time; about 250 million years ago these movements created Pangea
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endemic
species that are found in only one place in the world; islands generally have many endemic species; most island species are closely related to species form the nearest mainland or a neighboring island
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microevolution
the change in allele frequencies in a population over generations; evolution on its smallest scale; caused by (3) natural selection, genetic drift, AND gene flow
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discrete characters
characters that can be described in an either-or-basis; ex: purple or white flowers in pea plants; determined by a SINGLE gene locus with different alleles that produce distinct phenotypes
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quantitative characters
results from the influence of two or more genes on a single phenotypic character; resposible for most heritable variation; vary along a continuum within a population
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average heterozygosity
the average percent of loci that are heterozygous; estimated by looking at protein products of genes using electrophoresis; used to quantify gene variability (at the whole-gene level)
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nucleotide variability is measured...
by comparing the DNA squences of two individuals in a population and then averaging the data from many such comparisons
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geographic variation
differences in the genetic composition of seperate populations; ex: a cline, or graded change in a character along a geographic axis
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mutation
a change in the nuleotide sequence of an organism's DNA; ultimate source of new alleles
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RNA genome
what HIV has; it has a much higher mutation rate than DNA genome because of the lack of RNA repair mechanisms in host cells
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population
a group of individuals of the same species that live in the same area and interbreed, producing fertile offspring
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gene pool
all of the alleles for all the loci in all individuals of the population; a way to characterize a population's genetic makeup
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hardy-weinberg equilibrium
a gene pool where the frequencies of alleles and genotypes in a population remain constant from one generation to the next provided that ONLY Mendelian segregation and recombination of alleles are at work
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conditions for hardy-weinberg:
- 3 NO's!
- (1) no mutations (2) no natural selection (3) no gene flow
- 2 YES's!
- (1) random mating (2) large population size
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adaptive evolution
evolution that results in a better match between organisms and their environment; caused by natural selection when it consistantly favors some alleles over others
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genetic drift
chance events that cause allele frequencies to fluctuate unpredictably from one generation to the next, especially in small populations
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founder effect
when a few individuals become isolated from a larger population, this smaller group may establish a new population whose gene pool differs from the source population
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bottleneck effect
development of a new species caused by a severe drop in population size; by chance certain alleles may be over or underrepresented or event absent meaning there'll be low levels of genetic variation after
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genetic drift summary
(1) significant in small populations (2) causes allele frequencies to change at random (3) can lead to a loss of genetic variation within populations (4) can cause harmful alleles to become fixed
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gene flow
the transfer of alleles into or out of a population due to the movement of fertile individuals or their gametes; tends to reduce genetic differences between populations
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relative fitness
the contribution an individual makes to the gene pool of the next generation RELATIVE to the contributions of other individuals
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directional selection
occurs when conditions favor individuals exhibiting one extreme of a phenotypic range thereby shifting the frequency curve for the phenotypic character in one direction or the other; common when a population's environment changes or when members move to a different habitat
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disruptive selection
occurs when conditions favor individuals at both extremes of a phenotypic range over individuals with intermediate phenotypes
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stabilizing selection
acts against both extreme phenotypes and favors internediate variants; reduces variation; maintains status quo for a specific phenotypic character (ex: human baby birth weights)
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sexual selection
a form of natural selection in which individuals with certain inherited characteristics are more likely than other individials to obtain mates
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sexual dimorphism
marked differences between the two sexes in secondary sexual characteristics that AREN'T directly associated with reproduction or survival
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intrasexual selection
selection within the same sex, individuals of one sex compete directly for mates of the opposite sex; occurs among males (ex: peacocks)
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intersexual selection
individuals of one sex (usually females) are choosy in selecting their mates from the other sex; often times depends on the showiness of the males's appearance; also called mate choice
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balancing selection
occurs when natural selection maintains two or more forms in a population; inclues heterozygote advantage & frequency-dependent selection
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heterozygote advantage
occurs when organisms who are heterozygous at a particular locus have greater fitness than do both kinds of homozygotes (ex: fucking sick-cell anemia)
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frequency-dependent selection
when the fitness of a phenotype declines if it becomes too common in the population (ex: scale eating fish, left side/right side)
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neutral variation
when nucleotide differences in noncoding sequences appear to confer no selective advantage or disadvantage
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