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macroevolution
the pattern of evolution over large time scales
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3 examples of macroevolutionary change
- 1. origin of key biochemical processes such as photosynthesis
- 2. emergence of first terrestrial vertebrates
- 3. long-term impact of mass extinction on diversity of life
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earliest evidence of life
3.5 bya
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production of very simple cells thru 4 steps
- 1. abiotic (non-living) synthesis of small organic molecules: amino acids, nucleotides
- 2. joining of these molecules into macromolecules including proteins and nucleotides
- 3. packaging of these proteins into "protobionts," droplets w/membranes that maintained an internal chemistry different from that of their surroundings
- 4. the origin of self-replicating molecules that eventually made inheritance possible
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earth's early atmosphere was
reducing environment in which organic compounds formed from simple molecules
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Miller-Urey-type experiments demonstrate
that abiotic synthesis of organic molecules possible
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2 key properties of life
- accurate replication
- metabolism
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protobionts
collections of abiotically produced molecules surrounded by membrane-like structure
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first genetic material probably
RNA not DNA
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RNA
protein synthesis, enzyme-like catalyst functions (ribozyme)
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strata
sedimentary rock layers
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half life of C-14
5730 years
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stromatolites
layered rocks that form when certain prokaryotes bind thin films of sediment together
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cyanobacteria
oxygen-releasing, photosynthetic bacteria
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eukaryotic cells have
nuclear envelope, mitochondria, endoplasmic reticulum, cytoskeleton
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endosymbiosis
mitochondria and plastids were formerly small prokaryotes that began living within larger cells
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serial endosymbiosis
mitochondria evolved before plastids through sequence of endosymbiotic events
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the most widespread and diverse land animals are
arthropods (insects and spiders) and tetrapods
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key feature of nearly all prokaryotes is
the cell wall
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capsule
sticky layer of polysaccharide or protein, which allows prokaryote to adhere to substrate or to other individuals in a colony
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fimbrae (attachment pili)
hair-like protein appendages used to attach to substrate or one another
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taxis
movement towards/away from stimulus
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prokaryotes lack
membrane-bounded nucleus
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plasmid
small rings of separately replicating DNA
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ecndospore
when it copies its chromosome and surrounds it with tough wall. H2O removed, metabolism halts, remains dormant yet viable for a long time, hard to kill
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prokaryotes do/do not produce sexually
do not
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in transformation, genotype and possibly phenotype of a prokaryotic cell are
altered by uptake of foreign DNA from its surroundings
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in transduction,
bacteriophages carry bacterial genes from one host cell into another
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in conjugation,
genetic material is transferred b/w 2 bacterial cells that are temporarily joined
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the F factor
consists of about 25 genes, most required for production of sex pili
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R plasmids carry
resistance genes
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obligate aerobes
use O2 for cellular respiration
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obligate anaerobes are
poisoned by O2
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anaerobic respiration
substances other than O2 accept e- at downhill end of e- transport chains
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facultative anaerobes
use O2 if present but can also carry out anaerobic respiration of fermentation in an anaerobic environment
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nitrogen fixation
convert atmospheric nitrogen to NH3
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heterocytes can only do
N2 fixation
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biofilm
surface-coating colonies
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methanogens
use CO2 to oxidize H2 -> releasing methane as a waste product
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symbiosis
ecological relationship in which 2 species live in close contact w/one another
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parasitism
ecological relationship in which a parasite eats the cell contents, tissues, or body fluids of its host
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disease causing parasites
pathogens
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exotoxins
proteins secreted by certain bacteria and other organisms
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endotoxins
lipopolysaccharide components of outer membrane of gram negative bacteria
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bioremediation
the use of organisms to remove pollutants from air, soil, water
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viruses are/are not cells
are not
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capsid
protein shell enclosing the viral genome built from capsomeres
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viral envelopes
are derived from membranes of host cell, contain host cell phospholipids and membrane proteins
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bacteriophage
viruses that infect bacteria
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host range
specific variety of hosts that viruses can infect
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virulent phage
phage that reproduces only by lytic cycle
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restriction enzymes...
cut up DNA identified as foreign, restrict the ability of the phage to infect the bacterium
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lysogenic cycle
doesn't kill host
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prophage
when viral DNA integrated into bacterial chromosome
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retroviruses
RNA animal viruses w/most complicated reproductive cycles
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reverse transcriptase
transcribes RNA template into DNA, providing RNA -> DNA, which is opposite of usual direction
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3 ways that viruses burt on human scene
- 1. mutation of existing viruses
- 2. dissemination of a viral disease from a small, isolated human population
- 3. spread of existing viruses from other animals
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viral disease of plants spread 2 ways
- 1. horizontal transmission: infected from an external source
- 2. vertical transmission: infection by a parent
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viroid
circular RNA molecules that infect plants
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prion
infections proteins which appear to cause a number of degenerate brain diseases in various animal species.
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why prions are dangerous
long incubation time, virtually indestructible
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bioinformatics
application of computational methods to the storage and analysis of biological data
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physical map
distances b/w markers expressed by some physical measure, usually #base pairs along the DNA.
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physical map made by
cutting DNA of each chromosome into a # of restriction fragments then determining the original order of the fragments in the chromosomal DNA
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proteomics
systematic study of full protein sets encoded by genomes
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pseudogenes
former genes that have accumulated mutations over time and become non-functional
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repetitive DNA
consists of sequences that are present in multiple copies of the genome
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transposable elements
stretches of DNA that can move from one location to another within genome
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transposons
move within genome by means of a DNA intermediate
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retrotransposons
move by means of RNA intermediate that is a transcript of the retrotransposon DNA
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simple sequence DNA
contains many copies of tandemly repeated short sequences
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multigene families
collections of 2 or more identical/very similar genes
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homeobox
180-nucleotide sequence that specifies a 60-amino acid homeodomain in the encoded proteins
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zoospores
flagellated spores
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heteromorphic
the sporophytes and gametophytes are structurally different
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isomorphic
the sporophytes and gametophytes look similar
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Unikonta includes
opisthokont, amoebozoan
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Archaeplastida includes
- red algae
- green algae
- land plants
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Rhizaria includes
- forams
- radiolarians
- chlorarachniophytes
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excavata includes
- diplomonads
- parabasalids
- euglenozoa
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alveolata include
- dinoflagellates
- apicomplexans
- ciliates
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stramenopiles include
- diatoms
- golden algae
- brown algae
- oomycetes
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amoebozoans include
slime molds, gymnamoebas, entamoeba
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plasmodium
a single mass of cytoplasm that is undivided by plasma membranes and that contains many diploid nuclei
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producers
organisms that use energy from light (or inorganic chemicals) to convert carbon dioxide to organic compounds
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charophytes share 4 distinctive traits with land plants
- 1. rosette-shaped cellulose-synthesizing complexes
- 2. peroxisome enzymes
- 3. flagellated sperm
- 4. formation of a phragmoplast
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bryophytes do/do not have an extensive transport systems and include
do not
- liverworts
- hornworts
- mosses
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seedless vascular plants include
lycophytes and pterophytes
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grade
a collection of organisms that share a common level of biological organization or adaptation
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seed
an embryo packaged with a supply of nutrients inside a protective coat
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seed plants include
gymnosperms and angiosperms
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rhizoids
long, tubular cingle cells (in liverworts and hornworts) and filaments of cells (in mosses) that anchor gametophyte
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stomata
allow the exchange of CO2 and O2 between the outside air and the sporophyte interior, water evaporates from here
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xylem conducts
most of the water and minerals
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phloem distributes
sugars, amino acids, organic products
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most seedless vascular plants are homosporous/heterosporous
homosporous
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integument
envelops and protects the megasporangium
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sepals
usually green and enclose the flower before it opens
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stamens
produce microspores
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anther
where pollen is produced
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fungi do/do not ingest their food
do not
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hyphae
network of tiny filaments
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mycelium
interowven mass that infiltrates the material on which the fungus feeds
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septa
the hyphae are divided into cells by these cross-walls
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plasmogamy
the union of the cytoplasms of two parent mycelia
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