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homeotic (hox) genes
specifies the identity of a body part or segment during embryonic development.
ex. instruct the 3 thoracic segments (T1, T2, T3) each to develop a set of legs and the 2nd thoracic segment (T2) to develop wings
Hox gene clusters have been identified in a wide variety of animals with bilateral symmetry
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porifera
- sponges
- contain no tissues (Parazoa), but still has specialized cells
- still considered multicellular heterotrophic animals
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specialized cells of porifera
- porocytes: pores
- choanocytes: contains flagella to create current, also phagocytose food
- amoebocyte: shuttle food particles through the mesophyl
- spongocoel: main cavity
- osculum: exit, where current goes out
- spicules: silica or calcium carbonate skeleton
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Eumetazoa
all organisms that have tissue (will either be radial or bilateral)
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radial symmetry
- diploblastic
- necessary for slow or immobile organisms because they have a 360 degree response to their environment
- ancestral to bilateral symmetry
- Cnidarians all have radial symmetry.
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cnidarians
- radial symmetry
- diploblastic (2 embryonic tissue types): epiderm & endoderm
- can be polyp or medusa
- special features: cnidocytes (stinging cells on tentacles) and nematocysts (capture prey)
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Hydrozoa
- type of cnidarian
- polyp dominant (sessile) but also has a medusa stage (mobile)
- fire coral
- portuguese man-o-war (colony of polyp)
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Scyphozoa
- type of cnidarian
- true jellyfish
- exclusively medusa form
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Cubozoa
- type of cnidarian
- boxy jellyfish with extremely potent venom and highly developed eyes
- exclusively medusa form
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Anthozoa
- type of cnidarian
- include sea anemones & most corals
- strictly polyp form
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bilateral symmetry
- triploblastic (3 embryonic tissue types): endoderm, mesoderm, ectoderm
- coevolved with cephalization & movement
- includes protostomes & deuterostomes
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protostomes
- Blastopore develops into the mouth.
- spiral & determinate cleavage: Eary on, cells have a fixed destination so alteration to an embryo at an early stage is likely to be fatal.
- Mesoderm comes from solid masses of cells near the blastopore, which develop into the coelom (body cavity).
- includes mollusks, annelids, anthropods
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deuterostomes
- Blastopore develops into the anus
- indeterminate radial cleavage: Alteration to one cell early on is not likely to have an effect on the end form (totipotent)
- Folds of archenteron form coelom.
- includes echinoderms & chordates
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Lophotrochozoans
protostomes that have a lophophore (crown of ciliated tentacles around the mouth) and a larva phase during development
- flat worms (Platyhelminthes)
- Bryozoa
- Annelids
- Mollusks
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flat worms
- triploblastic & bilateral
- acoelomate (no body cavity): reason they are flat
- Turbellaria: free living (does not live inside an organisms's body)
- Monogeneans: parasitic on fish
- Trematodes: parasitic flukes
- Cestoidea: parasitic tapeworms
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ribbon worms (Nemerteans)
pseudocoelomates (false body cavity): only partially lined by mesoderm
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mollusks
- eucoelomates (full body cavities): complete gut
- has a mantle for gas exchange, a muscular food, only real shells, gills to filter feed (secondary for gas exchange)
- radula is a tongue-like structure
- include gastropods, cephalopods, bivalve, & chitons
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type of mollusk that has 8 separate shell plates (valves)
chiton (Polyplacophora)
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gastropods
- group of mollusks that include snails and slugs
- mostly hermaphrodites
- use radula to scrape algae off rocks
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cephalopods
- group of mollusks that include squid and octopi
- "head foot"
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bivalves
- group of mollusks that include clams and oysters
- has 2 shells (valves)
- incurrent and excurrent siphons
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characteristics of annelids
- eucoelomates (full body cavities)
- segmentation
- cutaneous respiration
- has complex organ systems, mouth, esophagus
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types of annelids
- Oligochaetes: earthworms
- Polychaetes: have parapodia (paired feet), live in oceans
- Hirudinea: leeches (parasitic)
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Ecdysozoans
- protostomes that shed or molt
- include nematodes & anthropods
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nematodes (roundworms)
- type of Ecdysozoan
- pseudocoelomate (false body cavity):
- has digestive & reproductive organs surrounded by a fluid-filled cavity
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arthropods
- group of Ecdysozoans that has jointed legs
- Hox genes important in segmented body plan giving rise to diverse appendages
- has an external skeleton made of chiton (polysaccharide)
- include chelicerates (spiders, scorpions, ticks), myriapods (centipedes & millipedes), crustaceans (lobsters, shrimp, crabs), and insects
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chelicerates
- arthropods that include spiders, scorpions, and horse-shoe crabs
- only anthropod that lack antennae
- has a cephalothorax
- pincerlike claws called chelicerae
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myriapods
- arthropods that include centipedes & millipedes
- many pairs of legs
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crustaceans
- arthropods that have branched legs
- has sensory antennae
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insects
- only arthropods to develop wings
- shed exoskeleton but cannot molt wings
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hexapods
- has 6 legs
- have a head, abdomen, and thorax
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echinoderms
- group of deuterostomes that include sea stars, sea urchins, sand dollars, and sea cucumbers
- has a water vascular system for respiration & movement
- has tube feet that allow them to crawl, sense their environment, capture food, and breathe
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Hemichordates have __
- a pharynx with pharyngeal slits
- a dorsal nerve cord
ex. acorn worms
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3 subphyla of chordates
- cephalochordates
- urochordates (tunicates)
- vertebrates (craniates)
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characteristics of chordates
- notochord (stiff rod of collagen)
- dorsal, hollow nerve cord
- muscular, post-anal tail
- pharyngeal slits or clefts
- myotomes
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urochordata
- tunicates (sea squirts)
- has a pharynx with numerous slits
- Water and wastes are expelled through an anal siphon.
- has a larva stage with chordate characteristics that are lost in the adult stage
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cephalochordates
- chordates that lack a well-differentiated head & has no cranium or vertebrate
- Instead, it has elongated bodies with a notochord that extends the length of the body from "head to toe"
ex. lancelets (amphioxus)
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agnatha
- jawless vertebrates
- includes myxini (hagfish) and lampreys
Hagfish has knotting behavior.
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gnathostomes
- jawed fish
- includes chondrichthyes & osteichthyes
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chondrichthyes
- cartilaginous fish
- has placoid scales (modified dermal scales)
- includes sharks and rays
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osteichthyes
- bony fish
- has dermal scales
- includes ray-finned fish and lobe-finned fish
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tetrapods
- amphibians and all amniotes (reptiles, mammals, birds)
- Ancestors may have been lobe-finned fish
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amnion
- desiccation-resistant shell
- can exchange gases while retaining water
- permit long development times
- allows organisms to be in dry terrestrial habitats
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amphibians
- do not have scales, absorb water directly through skin
- tetrapods but not amniotes
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reptiles
- scales made of alpha and beta keratin and formed from the epidermis
- Feathers of birds are modified epidermal scales that retain body heat.
- anapsids: no additional holes in the temporal region of the skull (tortoises)
- diapsids: 2 holes in the temporal region of the skull (crocodiles)
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characteristics of all mammals
- hair (permits retention of body heat)
- mammary glands
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prototherians
- monotremes
- lay eggs
- don't have nipples. Instead, the young drink milk secreted from pores in the skin of the mother's belly
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metatherians
- marsupials
- doesn't have a placenta so uses pouch
- The young are born at an early stage of development.
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eutherians
- placental mammals
- Placenta enables the offspring to be larger and more quickly independent when born.
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