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How do night time temperatures affect yield?
- High temps = Rd ^ = yield down
- Low temps = Rd down = yield ^
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Fick's Law
- Rate of diffusion ~ ∆concentration/∆distance
- ∆c/∆d
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affects of Still air v. windy air around corn plant on photosynthesis
- [CO2] @ 500m = 400ppm
- [CO2] @ leaf surface = Still 200ppm, Windy 400ppm
- ∆d gets much smaller = rate of diffusion
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What is N used for in plants
Protiens
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What is P used for in plants
- ATP
- NADPH
- PGA (phosphorylated sugars)
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What is K used in plants for
opening and closing stomata
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what is Mn used in plants
e- extraction from H2O
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Cu role in plants
in light rxn
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S role in plants
light rxn
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Fe role in plants
light rxn
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The formula
Ps-Rd = growth + stored reserves
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General Stored reserves
- No Ps when (hot, dry, cold)
- Survives on stored reserves
- Irreversible CHO Storage (seeds)
- Reversible Storage
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Where do Annuals store CHO
- Grasses: lower 1/3 of stem
- Dicots: Roots
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Where do Biennials store CHO
- Yr 1 - Veg devo and C stored
- Yr 2 seed production and no storage
- eg sweet clover = blood thinner
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Where Do Perennial Grasses store CHO
- 'stem base' (leaves/stem together, more leaves on younger) 3-4" above soil
- stems: may and june
- Rhizomes- below ground stem
- Stolons- above ground stem
- Corms- swollen internodes (timothy)
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Where do Perennial Dicots store CHO
- Taproots: carrots
- Stolons: white clover
- Rhizomes: not many (Kura clover)
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What kind of CHO compounds do plants store
- Sugars
- Polysaccharides (Starch, Fuctans)
- Lipids
- Protiens
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Stored plant sugar
4
- 4 kcal/g of material
- Sucrose: non-reducing sugar, inert
- C4 annuals are the main plants (corn, sorghum, millet)
- Also sugarcane and sugar beet
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Stored Plant starch characteristics
- Polymer of glucose
- 4 kcal/gram
- amylose 20%
- amylopectin 80%
- Not water soluable
- Stored in plastids called amyloplasts
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Amylose
- Starch
- linear glucose
- DP (degree of polymerization) = 3,000
- used to coat paper
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Amylopectin
- starch component
- branched glucose chain
- DP= 500,000
- why corn starch gels
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Plants that store starch
- C4 perennial grasses: zoysia, bermuda, miscanthus, switch grass, big blue stem
- Legumes: alfalfa, sweet clover, red clover, white clover
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Fructans
- Polymer of fructose (a polysaccharide)
- 4 kcal/g
- small DP= 3-300
- soluble in water
- cool season C3 (tall fescue, KY blue grass, orchard grass, wheat, barley, oats, rye, onion, Jerusalem artichoke, chicory)
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lipid storage in plants
- 9 kcal/g
- seeds
- small volume
- synthesis takes much more energy than CHO
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Plant proteins
- 4 kcal/g
- very expensive to synthesis
- N source- not N
- Vegetative storage protiens
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- clover canopy
- bad structure
- not all leaves involved
- gets bet out for light
- survives b/c N fixation
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- Grass canopy
- almost all tissues intercepting light
- taller than clover
- good canopy structure
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C4 temperature
- pretty bad lower than 10C/50F
- Optimum 30+/- 5
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C3 temperature
- Broad temp optimum
- 10-20C
- can still photosynthesis until ice forms
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both really decline when low
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Temperature affecting C3
- If no photorespiration and do really good at high temp
- Peak moves higher without
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When do plants fluoresce
Spell Fluorescence
- when light rxn screwed up
- fluoresce to get rid of energy
- atrizine stops e- passing = fluoresce and die
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around 20C how does temp ∆ dark respiration
- as temp goes up Rd rate doubles
- Q10=2
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H2O // CO2 exchange at leave ratio
more H2O lost than CO2 in
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Water Stress on Photosynthesis
- growth is stopped
- Ps not affected until mod stress
- drops off gradually as stomata close
- Stops at abrupt closure
- *Soybeans can get CO2 through cuticle
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Relationship btw leaf conductance and photosythsis
- As stomata open Ps increases
- If water limiting can partially close stomata- levels out Ps but cut water loss
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WUE
- water use efficiency
- kg H2O / kg Dry Matter
- alfalfa 1500:1
- Partial stomata closer helps this
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ratio of Ps to Rd when under stress
Rd >> Ps
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current CO2 concentration
- ~400 ppm
- winter goes up
- summer down
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Influence of [CO2] on leaf Ps of C3 & C4
- C4- no bump with increase
- C3- Photorespiration will go down @ 800 ppm CO2 will beat C4
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Influece of [CO2] on growth on C4 & C3
- C4 don't respond to increase
- C3 respond and grow more
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Ps of C4 and C3 with [O2] ∆
- C4: no impact
- C3: goes down with increase
- 2% O2 inhibits Photorespiration
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Influcence of wind speed and leaf size on boundary layer thickness
- big surface w/ slow wind = thick boundary layer
- thinner boundary layer as speeds go faster
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PEP carboxylase is how much of protein in C4s?
15%
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Influence of leaf [N] on Ps in C4 v. C3
- C4 responds better
- both increase
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C4 or C3 is more efficient on K use
C4
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Where are buds from tap root?
below the surface
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Non-structual CHO in alfalfa during season
Ps - Rd = growth and stored resese
- April: 0 - + = Up + down
- Late may: +++ - + = up + up
- August-cut: 0 - + = up + down
- need 155% of april to make it thorough winter
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N2 Fixation throughout season
- @ 100%
- cut = drop to 0
- for 2-3 wks fixation at zero
- then sharp increase
- N stored in taproots: gradually comes down then merges with fixation on the up
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Annuals stored reserve for grain growth
- straight line regardless of Ps
- Use stored reseves
- fill seed at all costs
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Cutting Alfalfa
- Cut close
- Not often
- will end up at the same level
- if every 2 wks- will deplete stored reserves and die
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Birdsfoot Trefoil
- Poor seedling vigor
- doesn't cause bloot
- effictively no stored reserves
- Needs to be cut high
- can be cut often
- photoperiods and cool = start storing
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Seasonal change in root N and protein in legumes
- Accumulates a lot in the fall (especially alfalfa)
- Goes down in spring for initial growth
- then up and down like in other charts
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N transfer values from storage for Rye grass
- 55% from stem base
- 47% roots
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N transfer values from storage for Alfalfa
- 41% from crown leaves and stem
- 78% from taproots
- 52% from lateral roots
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N transfer values from storage for White Clover
- 70% from stolons
- 56% from roots
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- Biennial
- year 1 mange like alfalfa (close, not often)
- Year 2 like Birdsfoot (often, not close)
- Don't want to cut in critical fall period
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Influence on defoliation on kernel dry wight in corn
- Ps is taken out of the model
- Rate is slightly reduced but still linear
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Influence of light intensity on restoration of nonstructural CHO in Alfalfa roots
- Takes longer in a low light environment to build back up
- doesn't ever make as much as high light
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Influence of temperatuer on CHO restoration in stem base of rye grass
- @ high temp: high Ps, low Rd = CHO up and storage happens
- @ low temp: drecase Ps, high Rd thus low CHO and growth
- NB: C3 fructan storage
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Influence of Stored reserves and leaf area on regrowth rate in Orchard grass
More important to have stored reserves than leaves
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How does N/P affect equation and lodging
- Ps up
- Rd ~
- growth ^^
- S.R. goes down
- 28% lodging
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How does NPK affect equation and lodging
- Ps ^^ (stomotal control)
- Rd ~
- Growth ^
- SR down little
- 1% lodging
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Affect of K on sugar mobilization in corn
If K added less of a drop in sugar
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Influence of N fert on leaf N, stubble N and yield
- More N =
- more leaf N
- less stubble N
- More yield
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Lodging Resistance
- Stem breakage = low CHO reserves in stem
- High K: prevents / limits lodging
- High N (P): can some times increase lodging
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N affect on equation
- Ps ^
- Rd ^ (more maintenance)
- Growth way up
- S.R. Down
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