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Geometric pop growth
applies to species that reproduce in synchrony during discrete time periods (seasonally). Occurs when pop changes in size by constant proportion from one time period to another.
- Nt+1 = ʎNt (Nt is pop size after t
- discrete time periods)
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- Nt = ʎt N0
- Ex: ʎ=2
- N0=100 individs
- What is predicted pop size at time t=20?
- N20 = 2.020(100) = 104,857,600 individs
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Exponential pop growth
- occurs when a continuously-reproducing pop
- changes size by a constant proportion at each instant in time.
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- dN/dt = rN
- r = exponential pop growth rate (intrinsic rate of increase)
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- → the rate of change in pop size at each instant in time.
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- (used to predict a pop size in the future, that
- grows continuously)
- Nt = N0ert
- Nt = pop size at time t.
- N0 = initial pop size (t=0)
- e is constant, base of ln(x)
- Geometric results in j-shaped points,
- exponential results in j-shaped curve
- Geometric and exponential growth overlap b/c equations are similar in form, ʎ is replaced by er
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Geometric/Exponential
How do ʎ and r affect pop size?
- ʎ = 1 or r = 0
- Nt+1= (1)Nt →Nt+1 = Nt
- dN/dt = 0 pop stays the same
- ʎ < 1 or r<0 pop decreases
- ʎ > 1 or r>0 pop increases
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Assumptions of geometric/exponential
growth
- Pop is closed (no migration)
- Constant birth and death rates (constant ʎ or r). Implies unlimited resources for growth
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Limits to pop growth
Density-independent factors – effects on birth&death rates are independent of pop density. same effects on small and large pops such as variation in weather, catastrophes
- Density-dependent factors - effects of birth, death, and dispersal (emigration) rates change with pop density. Intraspecific competition, predation.
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- ↑pop density = ↓birth rates. ↑death rates, ↑
- dispersal ↓pop growth (ʎ or r)
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Logistic population growth
- population increasing rapidly, then stabilizes
- at the carrying capacity.
- Carrying capacity (K) – max pop size that can be supported by the env.
- dN/dt = rate of change r = intrinsic
- rate of ↑ K= carrying capacity
- Suppose k = 1,000 When N is small (10), pop is growing at 99% of exponential growth
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- When N is close to k? (990). rN(0.01) When N exceeds k. dn/dt < 0. Pop=regulated.
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Assumptions of Logistic population
growth
- Closed population
- Constant carrying capacity
- Linear density dependence (growth rate decreases steadily as pop size approaches K)
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Nt+1 = ʎNt
Nt = ʎN0
geometric
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dN/dt = rN
Nt= N0ert
Exponential
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-/0
- amensalism
- elephants trample vegetation. Grass/shrubs are harmed, elephants are not. Penicillin and bacteria.
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Interspecific competition
- interaction b/w 2 species in which each is harmed by their shared use of a limiting
- resource.
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Types
of competition
- Exploitation competition – individs compete indirectly through their mutual effects on
- availability of shared, limited resources, ex plants depleting water.
- Sp. A (-)↔(-) Sp B
- (-) V (-)
- Limiting resource
- Interference competition – individs compete directly or access to a shared limiting resource.
- Ex hyena and vultures competing over a water buffalo, vine
- growing on tree.
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- Sp. A (-)↔(-) Sp B
- V
- resource
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Dominance-discovery tradeoff
another species goes in and takes over the resource.
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Competitive exclusion principle
2 species that use a limiting resource in the same way cannot coexist indefinitely.
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competitive coexistence
both species harmed, but neither die
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Resource partitioning
using resources in dif ways. Ex MacArthur (1958) warbler birds feed on insects in dif parts of the tree.
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types of exploitation
- Predation – kill and consume prey
- Herbivory – consume living plant tissues/fluids
- Parasitism – live in/on host, consume fluid
- Pathogens – cause diseases ex, malaria
- Parasitoids – kill host to complete life cycle (ex phorid fly-ant)
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Adaptations to exploitative interactions
- Defensive adaptations – when
- predators/herbivores exert strong selection pressure on prey and plants to evolve.
- Animals – large size, speed, body armor.
- Aposematism (warning coloration). Crypsis (camouflage), mimicry, behaviors.
- Plants – thorns, tough leaves, spines, toxic chemicals, chemicals that attract predators/parasitoids (tobacco and wasps killing caterpillars). Removal of plant tissues stimulates growth.
- Counter-adaptations – when prey and plants evolve to exert selection pressure back on predators and herbivores to evolve.
- Visual/chemical camouflage to sneak up on prey Tolerance or detoxification of chemical defenses (honey badger)
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types of mutualsim
- Facultative mutualism – each species
- can live w/o one another (ants and honeydew aphids)
- Obligate mutualism – neither species
- can exist without each other. (leafcutter ants and fungus)
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Producer
autotroph, primary producers measured in C or biomass
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Net primary production: gross primary production – respiration
Factors affecting npp
- Climate/avg temp
- Avg precipitation
- Nitrogen, phosphorous
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Consumer
- (heterotroph) – orgs that obtain energy by
- eating other organisms’ remains
Detritivores – eat dead organic matter
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Trophic levels
- 1st (primary producers, autotroph),
- 2nd (primary consumers, herbivores),
- 3rd (secondary consumers primary carnivore),
- 4th (tertiary consumers, secondary carnivores)
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Why pyramid shaped in terrestrial envs?
- Proportion of biomass at each trophic level not consumed
- Proportion of energy is lost in transfer
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Control of NPP
Bottom-up control
- resources that limlit NPP determine energy flow through an ecosys. Aquatic sys are limited by nutrients.
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- Silliman and Bertness(2002) manipulated snail density & found that grazing snails limit production of cordgrass. Removing the top predators (crabs ect) causes growth in snail pop, and decline in plant pop.
Ex of Trophic cascade – changes in abundance or species composition at one trophic level cause series of changes at other trophic level.
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Top-down control
energy flow is governed by rates of consumption at highest trophic levels
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