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Plants are different from animals:
- they are sessile
- they have indeterminate growth
- their support system uses organic, inert molecules
- they store different types of energy
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Root function:
- anchorage
- storage
- absorption
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taproot
- single main root with lateral roots growing off of it
- typical of eudicot
- adapted for deep soils
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fibrous roots
- adapted for shallow anchorage
- isodiametric
- may be adventitious - radical dies, roots emerge from stem itself
- typical of monocots
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modified root types:
- prop - comes out of root from stem (adventitious)
- storage
- strangling "aerial"
- buttress - supporting
- pneumatophores - in swampy areas, surface to "breathe" air
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stems/shoots are divided into...
- nodes - where leaves emerge, &
- internodes - stem between nodes
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2 types of buds:
- terminal bud - at top of stem, main bud
- axillary buds - above nodes, on the axis; termination of teminal bud will stimulate lateral growth
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modified stem types:
- stolon - grows horizonally and asexually; above ground
- bulbs - short, undergound; fleshy leaves surround it (ex: radish)
- tubers - grow at end of rhizomes (ex: potato)
- rhizomes - underground, horizontal; stem structures grow from it
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leaves on eudicots:
blades are connected to stem by a petiole
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leaves on monocots:
leaves emerge from stem base, forming a sheath
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modified leaf types:
- tendrils - demonstrate response to touch; may coil
- spines - nonphotosynthetic; defensive purposes (ex: cactus)
- storage - for water, nutrients (ex: jade, aloe)
- bracts - associated with flowers, colorful, main photosynthetic organ (ex: poinsetta)
- reproductive - serrated edges that grow additional leaves and roots; allots asexual reproduction
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parenchyma cells
- thin primary wall, not lignified
- useful for storage
- no chloroplasts, not much inside except for huge vacuoles
- isodiametric
- live at maturity
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chlorenchyma cells
- modified type of parenchyma cell with chloroplasts
- usually found in middle of leaves
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collenchyma cells
- unevenly thickened primary cell walls
- elongated cells
- provide support for actively growing cells
- living at maturity
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sclerenchyma cells
- tough, fibrous
- lignified secondary cell wall to help maintain shape (fibrous, barrel, star)
- dead at maturity; achieves correct shape/size then undergoes apoptosis
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general secondary cell wall characteristics:
- not very porous
- structure eliminates hyrdolic and turgor pressure
- interior to primary cell wall
- strengthened with lignin/cellulose
- thickness depends on what it differentiates into
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dermal tissues
- outer covering of leaves, stems, roots
- on leaves, may secrete cutin
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vascular tissues
- consist of xylem and phloem
- transportation tissues
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ground tissue
- neither dermal nor vascular
- provides bulk and support
- made of all cell types
- cortex - found exterior to vascular tissue
- pith - interior to vascular tissue
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meristematic tissue
- cells that undergo constant mitotic division
- consist of differentiating cells
- phloem points outside, xylem points inside
- apical meristems and lateral meristems
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xylem
- move water and dissolved inorganic nutrients
- modified sclerenchyma
- conducting elements are vessels and tracheids
- vessels: common in angios; wider, more efficient; have end plates - water can move easier from one element to another
- tracheids: in both angios and gymnos; narrow, spindle-shaped; assist in going through secondary cell walls
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phloem
- transport of photosynthate (dissolved sucrose)
- not necessarily a one-way flow
- modified parenchyma
- conducting elements are sieve cells (gymnos) and sieve-tube members (angios)
- sieve cells: living, but not a full compartment of organelles; albunious cells are helper cells
- sieve-tube members: missing many vital organs, but living; barrel-shaped for good efficiency; companion cells assist with organelle functions
- sieve plates - perforated plates; sap flows through these to adjacent cells
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apical meristems
- on root and shoot tips
- primary growth for herbaceous plants
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lateral meristems
- secondary growth in woody plants
- increase in girth, width
- meristems are the vascular and cork cambiums
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In roots, growth occurs in three overlapping zones of cells:
- zone of cell division - includes the root apical meristem and its derivatives; new cells are produced
- zone of maturation - most of the growth in roots; cells elongate and push root down further into soil; cell differentiation begins to become noticable
- zone of differentiation - cells become distinct cell types (at topmost part)
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vascular tissue organization in roots:
- monocots: parenchyma (pith) in center; core is surrounded by ring of xylem (inside) and ring of phloem (outside)
- eudicots: xylem (inside) and phloem (outside) in center core; stele is the vasculer cylinder appearing as a lobed core
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Monocot root tissue organization (outside to inside):
epidermis --> cortex --> endodermis --> pericycle --> vascular cylinder (phloem, xylem)
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Eudicot root tissue organization (outside to inside):
epidermis --> cortex --> endodermis --> pericycle --> phloem --> xylem --> pith(core of parenchyma cells)
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endodermis
- special layer of cells with super thick walls that are water proof and lignified
- selective barrier for stele
- modified parenchyma
- innermost layer of cortex
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primary growth of shoots:
- leaves develop from leaf primordials - projections along the side of the apical meristem
- shoot elongation is due to lengthening of internode cells just below the shoot tip
- branching arises from activation of axillary buds
- intercalary meristems allow damaged leaves to rapidly regrow
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Monocot stem tissue organization (outside to inside):
epidermis --> [maybe] collenchyma cells --> cortex --> vascular bundle (sclerenchyma --> phloem --> xylem) --> pith
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Eudicot stem tissue organization (outside to inside):
epidermis --> collenchyma cells --> scattered vascular bundles within mass of ground tissue
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Leaf tissue organization (top to bottom):
upper epidermis (cuticle --> layer of dermal tissue interrupted by stomata and guard cells) --> palisade mesophyll (elongated chlorenchyma cells) --> vascular tissue (surrounded by bundle-sheath cells) --> spongy mesophyll (contains air spaces) --> lower epidermis
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secondary growth
- aid from lateral meristems (cambiums)
- only in woody plants
- builds on primary growth plan
- typical in eudicots and gymnos
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cork cambium
- provides protective layer for stem
- cork on the exteriror
- corticle cells on the interior
- entire complex called bark (which also includes secondary phloem)
- develops within cortex
- replaces epidermis
- periderm (cork cambium) secretes suberin into cell walls that makes the bark water proof
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vascular cambium
- produces secondary phloem to exterior and secondary xylem to interior
- youngest tissue closer to vascular cambium, older tissue towards center of trunk
- phloem layers are much narrower
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bark
everything outside of vascular cambium : periderm (cork cambium and cork) and secondary phloem
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lenticels
loosely arranged cork cells that oxygen can diffuse through
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girdling a tree
taking an entire ring of bark off of the tree can kill it because of secondary phloem removal
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