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How do we assess global diversity? What is the ultimate source of biodiversity?
- Changes or mutations (alternations)in the genetic code (DNA/RNA)
- In both coding and non-coding regions
- Accumulation of mutations of non-coding regions also tells us the evolutionary relationship of organisms
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What are the 3-levels that we measure genetic variations within a species?
- 1.) At an individual level (some individuals in a population maybe fixed for two dominate alleles that have no variation at the gene locus)
- 2.) Within population variations (One individual might have a genotype of AA, but there might be several individuals within the population with a little "a" allele
- 3.) Between population variations (Different populations become adapted locally and have genetic divergence)
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Why is variation important in populations?
Populations that tend to be more genetically diverse, on average, are considered to be a healthier population
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Non-coding regions
introns
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Heterosis (Heterozygous advantage or hybrid vigor)
Heterozygous offspring have a higher survival and fitness compared to homozygous parents
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Fitness (biological fitness)
relative reproductive output of an individual in a population compared to everyone else
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In conservation programs, biodiversity and heterosis suggests:
that we try to maintain or maximize genetic diversity within a population and with individuals
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Why is DNA a more sensitive tool compared to using proteins?
- DNA codes for the same amino acids
- DNA is a more sensitive tool because a lot of silent mutations don't affect proteins that are produced from the genetic code or sequence
- Change in genetic sequence which increases the genetic variation at the DNA level, but wouldn't result in an increase variation at the protein level
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When a population get bigger in size, What happens to the amount of heterzygousity or genetic variation in a population?
It increases
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Rates of genetic variation
- Tends to increase with bigger population size, but at different rate for different organisms
- Some taxa evolve at different rates than other taxa
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Correlation Co-efficient (r)
tells the strength of a linear relationship
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If two variables are plotted
Y=%
X= population size
If X increases and Y increases, what is the range of (r)
- r= 0 to +1 (positive correlation)
- 0 means no correlation and +1 means perfect correclation
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If two variables are plotted
Y=%
X=population size
If X increases and Y decreases, what is the range for (r)
- r= -1 to +1
- +1 means all the points fall in a perfect straight line on a positive slope
- -1 means all the points fall in a perfect straight line on a negative slope
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What % of the relationship of y is explained by the variation of x?
(If the co-efficient is squared and multiplied by 100, it turns it into a %)
- Example: r= .94
- (.94)2(100) = 88%
- **12% is something else than population size
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Cheetahs are an exception to the rule
- Cheetahs are highly inbreed
- Very little genetic diversity between individuals (inbreeding and inbreeding depression can occur)
- Have a high infant mortality rate (doesn't matter if inbred or outbred)
- Genetically weak species
- Probably went through a genetic bottleneck at some point in their history
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Inbreeding
mating between close relatives
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Inbreeding depression
loss in vigor and fitness due to an increase in homozygousity
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Organisms that are highly inbred do not have a lot of genetic variation because they have a very limited gene pool
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Why does inbreeding depression occur when close relatives reproduce?
- Heterosis = less genetic variation (reduces survival and fitness)
- Genetic mutations (higher in our family line than in the general public)
- Recessive alleles as pairs
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What causes populations to vary genetically?
- 1.) Natural selection (some genotypes/phenotypes are favored in some environments over other enviroments; different selective pressures)
- Example of frog: webbing on feet
- 2.) Neutral evolution mechanisms
- (genetic drift, founder's effect, and bottlenecks)
- Only effects neutral alleles, which does not directly affect the survival or fitness of an organism
- Genetic drift is usually for in small populations
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Neutral alleles
are alleles that do not directly affect the survival or fitness of an organism
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What two factors do modern biologists believe drive the evolution of populations?
Natural selection and neutral evolution mechanisms
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Genetic drift
random loss/fixation of alleles in populations (has nothing to do with selective advantages)
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What are some possible explainations of why two populations would not be genetically divergent from one another?
- Gene flow (1st answer you should think of)
- Same environment (where the same genotypes/phenotypes are favored)
- Separated recently (suggests that the two populations were recently separated and have not had the time to accumulate genetic differences)
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How do we assess biodiversity of a community or habitat?
- Simplest measure is species richness (S)
- ** it is not a very useful metric because it does not utilize/report the relative abundance of individuals in each species
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Species richness (S)
# of species trait that occur in a given community, habitat, or ecosystem
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How do we measure abundance?
- 1.) Count individuals and convert our counts into proportions
- Example: 90 out of 100= .9
- 2.) Use fractional amount of area that each species covers or occupies
- Proportion of area covered by the species (like plants)
- Example: Plant "A" occupied 50% of the area covered = .5
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Better measures for biodiversity
Diversity indices take into account the relative abundance of each species besides just counting to see if one species is available/present or not
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2-types of diversity indices
- 1.) Dominance indices
- Example: Simpson's dominant index
- 2.) Indices for sensitive to rare species
- Example: Shannon index
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Dominance indices
- They give much more influence to the index to common (= dominant) species
- Good when you have a lot of dominant individuals in a community
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Simpson's dominant index
- delta with subscript S = the sum of 1/pi squared
- where pi is the proportion of individual species i
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Indices for sensitive to rare species
- Good for conservation programs because we are dealing with rare species
- Gives some influence to rare species when you calculate the index
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Shannon Index
H'=-sum of (pi(lnpi))
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Indicator species
usually animals or plants; organisms that are used as a surrogate to assess the conditions (health) of an ecosystem
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How do you chose an indicator species?
- Depends on the environment
- Examples:
- 1.) Amphibians (frogs and salamanders) = fresh water habitat
- They are tied to water for cutaneous respiration and reproduction
- 2.) Lichens (obligate mutualism between fungus and algae) = air quality
- 3.) Spotted owl = old growth forests
- (they are monitored because they are endangered and because they are used as indicator species)
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2- ways to use indicator species
- 1.) Use endemic species and monitor for changes in their distribution and abundance (also health)
- Looking for population declines
- 2.) Use invasive species as an indicator of environmental change
- Do not naturally occur in they community, but have showed up in the community due to enviromental changes
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Endemic species
species found in a community naturally and nowhere else
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Invasive species
species that are not naturally occuring in a community
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Halophytes
organisms that like to live in salty conditions
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Problems with indicator species
- 1.) Don't know the effect of the indicator species on the other organisms in ecosystem
- 2.) Extrapolation to other species may be invalid because each has its own requirements
- 3.) Invalid monitoring schedule (don't follow the species long enough)
- 4.) Biased choice of indicator species; often verts and relatively large (Higher in trophic levels and are not sensitive to early changes in the environment)
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Keystone species
- European Honeybees
- Gopher tortoises
- African elephants
- Sea otters
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50/500 rule (maintain population)
- n= 50 (prevent inbreeding depression)
- n= 500 (prevent genetic drift)
- Less than 50= populations are endanger of both inbreeding depression and genetic drift
- Less than 500= population protected from inbreeding depression, but loose alleles due to genetic drift
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Problems when establishing a conservation reserve
- 1.) Species list (gives a list, but not the abundance)
- 2.) Do not know the species diversity
- 3.) Many organisms live in ecosystems with poorly defined boundaries (How do we know where to set-up boundaries)
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Biome
- Large-scale community that can be indentified based on the dominant vegetational types
- Plants are important and define a biome and what kind of biome in a given area
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2- most important factors that determine a biome
- 1.) annual average temp
- 2.) annual average ppt
- Temp in. and ppt de. = desert
- Temp de. and ppt de. = tundra (cold des.)
- Temp. in. and ppt in. = t.r.f
- Temp intermed. and ppt in. = temp. rainforest
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Biodiversity
- 1.5 to 2.0 million extant species been described by science
- 300,000 extinct species
- Estimated 10 to 50 extant species
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Why are estimates so variable?
- Depends on species concept used
- Biased; based on species that affect humans
- Tend to describe larger species
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Some groups underrepresented
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1975 Strong and Levin
- Fungal species went from 80,000 to 1.5 million species
- Stated that there are 6 unique fungal species per plant species (6X 250,000)
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Earth 4.2 to 4.5 billion years old
1st 100 million years too hot to support life
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Oldest fossils approx. 3.8 to 4.0 billion years old
- Prokaryotes
- No membrane bound organelles
- Non-membrane bound nucleus
- DNA floats around freely in the nucleod
- Few organelles (mitochondria)
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Organisms in higher cells that were swallowed up that were free-living
- Mitochondria
- Cholorplasts
- Centrials
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Evidence for free-living organisms
- Double membrane
- Own DNA
- The divide independently of nucleus
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Endosymbiotic theory
some of our complex eur. cells which have organelles became part of cell because larger cells swallowed up and did not digest.... reached an agreement
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Riapaian
a long banks of fresh flowing water
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